Tom McGrath
Skipper (voice)

Chris Miller
Kowalski (voice)

Christopher Knights
Private (voice)

Conrad Vernon
Rico (voice)

John Malkovich
Dave (voice)

Benedict Cumberbatch
Classified (voice)

Ken Jeong
Short Fuse (voice)

Annet Mahendru
Eva (voice)

Peter Stormare
Corporal (voice)

Andy Richter
Mort (voice)

Danny Jacobs
King Julien (voice)

Werner Herzog
Documentary Filmmaker (voice)

Sean Charmatz
Cricket (voice)

Billy Eichner
New York Reporter (voice)

Stephen Kearin
Pilot / Aquarium Employee (voice)

Ava Acres
Girl at Zoo (voice)

Kelly Cooney Cilella
Mermaid Penguin (voice)

Susan Fitzer
Antarctic Penguin (voice)

Chris Sanders
Antarctic Penguin (voice)

Emily Nordwind
Antarctic Penguin (voice)

Mike Mitchell
Antarctic Penguin (voice)

Hope Levy
Antarctic Penguin / Woman at Aquarium (voice)

Walt Dohrn
Antarctic Penguin (voice)

Jim Pirri
Gondolier (voice)

Elizabeth Pan
Shanghai TV Reporter (voice)

Jeff Fischer
Security System Voice (voice)

Stephen Apostolina
Penguin Prisoner (voice)

Al Rodrigo
Penguin Fan (voice)

Steve Alterman
Aquarium Announcer (voice)

Carter Hastings
Boy at Aquarium (voice)

Cullen McCarthy
Boy at Zoo (voice)

Nicholas Guest
Flight Attendant (voice)

Adriano Aragon
Angry Man (voice)

Lynnanne Zager
Computer Voice (voice)

Angie Wu
Girl with Snow Globe (voice)